Pretty sure they are just the start/stop points of lines(seams). Other than printing in vase mode they are nearly impossible to totally get rid of due to the nature of FDM printing. Things like coast, wipe, pressure advance will all help but when you have molten plastic being laid down in lines that start and stop there's not much that can be done.
Best thing you can do is properly tune your extruder, filament flow and work with pressure advance.